The original Monkey Ball characters of Aiai, Meemee, Gongon & Baby are back. They are joined by a host of new characters, including a Prince and Princess, who are being kept apart by a feud between their kingdoms. The couple run away to Jungle Island where they enlist the help of the Super Monkey Ball gang, to unite the feuding Monkey Kingdoms and defeat the Naysayers who have sucked all the joy from the Monkey Ball world.